Red fluorescent proteins (DsRed) discovered in anthozoan corals. (19) Leads to discovery of many new fluorescent proteins and chromoproteins. GFP tolerates insertion of entire proteins in certain locations and can be circularly permuted.

Reference: MV Matz, AF Fradkov, YA Labas, AP Savitisky, AG Zaraisky, ML Markelov, SA Lukyanov: Fluorescent proteins from nonbioluminescent Anthozoa species. Nature Biotech. 17 (1999) 969-73.